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Description:
The exact taxonomic status of this arthropod is in dispute. It has
been termed a "trilobitomorph" by some researchers, showing
their beliefs in its close affinity with the trilobites. While the
Trilobitomorpha was listed in the Treatise, most now consider that
this subphylum is invalid, a catchall much like some of the dustbin
terms used by Walcott for a number of the Burgess Shale fauna. Whittington
termed it a "soft-bodied trilobite"' but that belief is
not supported at present. Their similarity in appearance to the
Agnoistida is purely a result of convergence. Two large groupings
of the Paleozoic arthropods are currently in favor: the Crustaceomorpha
(which includes Waptia) and the Arachnomorpha, dominated by the
trilobites. The Naraoiidae are arachnomorphs and include Misszhouia
and Naraoia.
Misszhouia longicaudata
was initially placed within Naraoia, but was given its own genus
based upon differences between it an Naraoia copacta of the Burgess
Shale. The species is one found in several locations within Yunnan
Province, but this one is from the most famous location of all,
Maotianshan, site of the discovery of the Chegjiang Biota by Hou
Xian-guang in 1984. Indeed, it is this taxon which was first discovered
at Maotianshan to start the entire cycle of research.
The diversity
of soft-tissue fossils at Jengjiang is astonishing: algae, medusiforms,
sponges, priapulids, annelid-like worms, echinoderms, arthropods
(including trilobites), hemichordates, chordates, and the first
agnathan fish make up just a small fraction of the total. Numerous
problematic forms are known as well, some of which may have represented
failed attempts at diversity that did not persist to the present
day.
This specimen
shows many of the classic features of the taxon, even the diverticula
and the central gut are evidence, as is the soft cuticle that betrays
the outline of the body. Even if it is not a true trilobite, it
is sure to be a coveted addition to any collection, one of historical
significance due to its ties with the discovery of the Chengjiang
Biota.
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