Darwin
both pondered and worried about the paucity of fossils found
that dated before the Cambrian. More than a century later we
now know that arthropods lacking external exoskeletons walked
in the shallows, on the beaches, and ultimately made their appearance
on land. Darwin would today be gratified that science has determined
that these transitional creatures had left there traces, if
only footprints in the sand, substantiating there overarching
of the Precambrian/Cambrian boundary of time. This fossil is
such evidence, coming from the history-making Krukowski
quarry in central Wisconsin that has only recently come
under intense study by Paleontologists. New publications are
forthcoming, adding to several in the last year (e.g., see Stranded
on a Late Cambrian
shoreline:
Medusae from central Wisconsin, by Hagadorn, et. al., describing
a diverse fauna that apparently walked or floated upon an ancient
Cambrian shoreline.
The
Diplichnites shown to the right is unique owing to the obvious
large size of the animal, since the sides of the tracks are
some 2 inches apart. Note that this large specimen is the convex
half of the fossil (called hyporelief by Ichnologists). This
particular plate measures 775 by 140 (30 1/2 by 5 1/2 inches),
and a yardstick is shown to give perspective.
Popular
conjecture is that Diplichnites is the trace of a wandering
Myriapoda, which brings to mind an ancient centipede (Chilopoda).
Given the spacing of these footprints, the centipede that made
the tracks of the fossil to the right would have been an awesome
predator - perhaps the terror of the Cambrian beach some 1/2
billion years ago.