Probably
the most famous Lagerstätte in the world is that
of the Solnhofen limestone deposits in the state of Bavaria
in Germany. The
amazing diversity of fossil organisms, often exquisitely
preserved, provides a glimpse into the past, 155 million
years ago. The
most famous among them are the eight fossils of the first
bird known in detail, Archaeopteryx. The first Archaeopteryx
fossil was discovered in 1860, a single feather. The following
year a very complete example was found, which was ultimately
sold to the British Museum. The next specimen was found
sixteen
years later, and is known as the Berlin specimen. It is considered
by some the most valuable fossil in the world today. With
its combination of characteristics of reptilian-like teeth
and tail with the feathers of a bird, it was an ideal "missing
link" the supporters of Charles Darwin could point to
as proof of his theory of evolution, first published in 1859.
In the following 140 plus years since the first bird was found,
only six more have been added to the ranks of examples.
The
discovery of the wonderful fossils of Solnhofen may be
attributed
to the longtime uses to which the "plattenkalk"
has been used. This is a German word that seems more appropriate
than the English "platy limestone". Plattenkalk
was used as early as the Stone Age for making drawings and
colored murals due to its softness. The flat, regularly shaped
material was suitable for paving roads and building walls,
something done by the ancient Romans who for a time held
sway
over the region. In the Middle Ages, the stone was used as
floor and roofing material. The mosaic floor of the church
of Hagia Sofia in Istanbul was made of limestone from the
region. Artisans also used the material in the making of
bas-relief
sculptures and headstones. A decisive turning point in the
history of the area was the discovery in 1798 by Alois Senefelder
that the fine-grained material could be employed in lithography,
a use that caused quarrying to skyrocket. Although the heyday
of lithography only lasted a hundred years or so, it was
the
single most important impetus in the discovery of fossils.
The
fossils have always been prized by local residents because
of their beauty. As those most intimately associated with
the limestone, the quarrymen were obviously those who were
responsible for their discovery. At first, quarry owners allowed
the workers to keep the fossils they found, but as interest
in them (and consequently, value) grew, this practice stopped.
The variety and number of fossils known is deceptive. The
occurrence of fossils is quite low. Indeed, a worker can quarry
for an entire day and find not a single one. The hundreds
of years of quarrying are what make them seem so apparently
common.
The
sheets of limestone are so regular that we can only conclude
that they were laid down in a calm environment. The deposits
were evidently laid down under a stable body of water that
had some connection to the Tethys Sea. Sponges and corals
grew on rises in this sea, forming lagoons. In fact, remnants
of a coral reef can be found in the area to the south of Solnhofen.
The region must have been near land, however, due to the discovery
of insects such as wonderfully-preserved dragonflies. Assuming
that Jurassic coral reefs grew in modes similar to those of
today, the surface of the reefs would have been only 10 meters
or so under water. The maximum depth has been estimated by
some to have been 60 meters (200 feet).
These
isolated lagoons would have been quite stagnant due to little
exchange of water with the sea. Anoxic conditions would have
been ideal for preventing destruction of organisms that found
their way into the lagoons. Some evidently survived for short
periods. One of the most famous examples of this are the horseshoe
crab "death spirals' that exist in which a spiral trackway
has been preserved with the defunct arthropod in the center,
presumably preserving its last efforts at survival. The theory
most often proposed for the toxicity of the waters has been
that of hypersalinity, the excessive concentration of salt.
If the area were hot and dry, with little runoff from the
land to the north, conditions would have been ideal for promoting
excessive evaporation of water with concomitant increase in
the salinity in the lagoons. The dense brine would collect
in the bottom of the pools, excluding most life, as sensitivity
to even minute changes in density has been seen in many marine
organisms. Once an organism had been washed into the lagoon
by the action of a storm, it would quickly succumb to the
toxic conditions that existed within. The hypersaline, anoxic
floor was ideal for the preservation of the body, often even
leaving evidence of soft tissues.
Storms
would have often brought in a suspension of finely-dispersed
lime. Such storms would have brought in the amazing diversity
of life we know from the region: pterosaurs, dinosaurs like
Compsognathus and Archaeopteryx, dragonflies, and other insects,
fish, turtles, crinoids, starfish, jellyfish, ammonites, worms,
plants, and many, many more. The number of species found exceeds
500. The limestone deposits of Solnhofen open a door closed
to us over 150 million years ago, affording us a look at a
wonderful diversity of life from both the land and the sea
of the Jurassic.